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21. Syngonium harlingianum Croat,
sp. nov. TYPE: Ecuador, Pastaza, Mera, Pantanos, 1100 m, Harling 11064
Planta terristris, caudex ca. 1 cm diam.; lamina simplex trilobata,
lobi conjuncti, lobus medius 7.5-10.5 cm longus, 2.7-6 cm latus, lobi
laterales inaequilaterae conspicuae auriculatae. Inflorescentia solitaria,
9.5 cm longa, spatha 11.5 cm longa, tubo 4.5 cm longo, ca. 2 cm lato,
viridi, lamina luteola, aeuminata; spadix 8.5 cm longa, parte feminea
spadicis ca. 2 cm longa.
Terrestrial. Adult plants with stems ca. 1 cm diam.; internodes 1-2
cm long near the apex; petioles 12-14 cm long, sheathed ca. 2/3 their
length, the free portion subterete, ca. 2 mm diam., the sheath merging
almost imperceptibly with the petiole at its apex; blades 3-lobed,
drying yellowish green, the median lobe oblong-elliptic, acuminate
to obtuse at the apex, narrowed at the base and confluent with the
lateral lobes, 7.5-10.5 cm long, 2.7-6 cm wide, the lateral lobes
markedly inequilateral (the inner part Vi or less as wide as the outer
part), more or less elliptic, obtuse to acute at the apex, 5.5-8 cm
long, 2-4 cm wide, rounded or more commonly auriculate on the outer
edge at base, the midrib naked for 5-10 mm, the auricles to 2.5 cm
long, rounded or oblong, not at all constricted at the base, the surfaces
drying matte, yellowish green, slightly paler beneath; midrib drying
flat on upper surface, weakly raised beneath with obscure scabrid
primary lateral veins 2-4 pairs; tertiary and quaternary veins distinctly
primary collective vein arising from the first or second primary lateral
vein, 2-4 mm from the margin.
Inflorescences solitary; peduncle 9.5 cm long, ca. 3
mm diam. (dried); spathe 11.5 cm long; spathe tube green, narrowly
ellipsoid, 4.5 cm long, ca. 2 cm diam.; spathe blade narrowly elliptic,
membranous, yellowish, acuminate at the apex; spadix 8.5 cm long;
pistillate portion of the spadix ca. 2 cm long, ca. 5 mm diam., the
flowers contiguous, ca. 1 mm diam.; sterile stam-inate flowers irregularly
elongated in the direction of the axis, 3-3.5 mm long in direction
of the axis, 1-1.6 mm wide perpendicular to axis; staminate portion
of the spadix cylindroid, narrowly tapered toward the apex, drying
ca. 1 cm diam., the flowers mostly rhombic, the apex truncate, matte,
the margin crenate, not at all incised between the anthers.
Infructescences not seen. Fig. 44.
DISTRIBUTION: Syngonium harlingianum is known only from the
type collection and is named in honor of its collector, Dr. Gunnar
Harling, one of the coauthors of the Flora of Ecuador.
The species is not closely related to any other species nor is it
confused with any other species. It is characterized by its small,
3-lobed leaves with confluent leaflets. It was described by its collector
as being terrestrial, an unusual condition in the genus. While it
is not unusual to find juvenile and pre-adult leaves on the ground,
I have never found adult plants in other than an epiphytic or hemiepi-phytic
The type collection was in flower in late February.
ECUADOR: PASTAZA: Mera, Harling 11064 (GB).