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Syngonium sparreorum Croat, sp. nov.
TYPE: Ecuador, Cotopaxi, Rio Gua-para, ca. 20 km NW of El Corazon,
monsoon forest, ca. 250 m, Sparre 17193 (S, holotype).
Caudex 1-1.5 cm diam.; lamina trisecta aut 5-pedatisecta; petiolus
29-47 cm longus, ad 1/3-2/3 longitudinalis usque anguste vaginatus;
lamina elliptica ad obovata-elliptica, segmentum medium equalis, segmenta
lateralia inaequalia leviter, venis lateralibus utroque latere 10-12,
fere rectangula-ribus. Inflorescentiae 2; pedunculus 7-10 cm longus,
gracilis; spathae tubus 5-6 cm longus, oblongus-ovoideus, 1.5-2 cm
diam.; spathae lamina 7.5-8.5 cm longa, ovata, apice cuspidate; spadix
parte feminea 10-12 mm longa, parte mascula spadicis 6-7.5 cm longa;
infructescentia pendula, spathae tubo atropurpureo; spadix ad 9.5
cm longa; semina atera, ca. 7 mm longa.
Hemiepiphytic vine; stem 1-1.5 cm diam., drying pale brown; internodes
elongate, 6-7 cm long near the apex; petioles 29-47 cm long, narrowly
sheathed Vi-2/3 their length, to 7.5-10 cm from the apex, the apex
of the sheath slightly free from the petiole; blades trisect to 5-pedatisect,
the leaflets distinctly free; median leaflet equilateral, elliptic
to obovate-elliptic, abruptly acuminate at the apex, cuneate to acute
at the base, 17-28 cm long, 9.5-12 cm wide, the leaf tissue decurrent
to near the base, the lateral leaflets slightly inequilateral, obtuse
to acuminate at the apex, 14-25 cm long, 5.3-9 cm wide, the base narrowly
acute on the inner side, obtuse to rounded or conspicuously auriculate
on the outer side, the auricles conspicuously constricted at their
base; petiolules 7-17 mm long, sometimes free; outermost leaflets
nearly oblong to ovate-oblong, 8-10 cm long, 2.7-7 cm wide, narrowly
rounded to acute at the apex, the blade surfaces drying yellowish
brown; primary lateral veins mostly 10-12 pairs, mostly joining the
midrib at a nearly 90° angle, curved to the collective vein; collective
vein 3-10 mm from the margin, almost straight or conspicuously loop-connecting
with the primary lateral veins; reticulate veins very fine and close
but clearly visible on dried material.
Inflorescences 2 in uppermost leaf axil, erect; prophylls
ca. 15 cm long, obtuse at the apex, apparently caducous; peduncles
7-10 cm long, slender, 2-4 mm diam. (dried); spathe tube 5-6 cm long,
oblong-ovoid, 1.5-2 cm diam.; spathe blade 7.5-8.5 cm long, ovate
when open, cuspidate at the apex, thinly coriaceous, drying brown,
soon caducous; pistillate portion of the spadix 10-12 mm long, 5 mm
diam. (dried), the flowers ca. 1.5 mm diam., in 5 or 6 spirals, the
stigmas raised, ca. 1 mm diam.; staminate portion of the spadix 6-7.5
cm long, clavate, ca. 1 cm diam. in the widest part, conspicuously
constricted just above the sterile staminate flowers, the fertile
staminate flowers ca. 3 mm long, rounded to irregularly 4-6-sided,
the apex of the synandrium truncate, entire, the margins minutely
crenulate; sterile staminate portion of the spadix ca. 14 mm long,
ca. 8 mm diam. the sterile staminate flowers ca. 2.5 mm long, 1.5-3
mm diam., mostly free from one another, rarely fused.
Infructescences pendent; peduncle to 13 cm long, to
7 mm diam.; spathe tube oblong-ellipsoid, gradually tapered to both
ends, to 13 cm long and 5 cm diam., apparently dark violet purple;
fruiting spadix oblong-ovoid, to 9.5 cm long on the front side, 6.5
cm long on the back side, ca. 4.5 cm diam.; seeds black, oblong-ovoid
ca. 7 mm long. Figs. 71, 72.
DISTRIBUTION: The species is known only from the type locality on
the Pacific slope of northcentral Ecuador.
It is named in honor of the co-author of the Flora of Ecuador, Baron
Benkt Sparre, who has made the only collections of the species.
Syngonium sparreorum is probably closest to S.
gentryanum from Huanuco Department, Peru, but differs
in being 5-pedatisect in part and in having the primary lateral veins
of the leaflets more numerous and joining the midrib at a nearly 90°
Flowers and immature fruits were found in June.
ECUADOR: COTOPAXI: Rio Guapara NW of El Corazon, 250 m, Sparre 17096,
17101, 17193, 17212, 17297 (S).