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  Re: [Aroid-l] Alocasia macrorrhizos and A. cucullata
From: Peter Boyce <phymatarum at googlemail.com> on 2009.09.06 at 06:56:11(19945)
Hi LariAnn,

To re-iterate my reply to Tony, Certainly 'species' such as A. macrorrizos
and A. cucullata bend the boundaries a lot. What is of course interesting is
that A. macrorrhizos (notwithstanding its doubtful 'pure' species status) is
definitely related (and here we are talking molecularly) to some
unquestionably 'good' species, such as A. portei and A. flabellifer, which
poses even more difficulties. It is also problematic to talk about
utilization of cultivars, especially those that are selections of what may
themselves be cultivars, albeit so long-standing that they have effectively
stabilized and function as species, even to the extent that they have lost
the ability to hybridize with other elements of what was once a single gene
pool.

The bottom line is that, at present, we can only be sure that A.
macrorrhizos and A. cucculata are NEVER found away from human disturbance in
'habitat' and furthermore, away from the attentions of horticulturists are
remarkable morphologically stable.

As a final thought on this, it is also important to remember that species
framework, and the interspecific crossing is often in nature not just a
matter of 'incompatibility'. Distribution, flowering time even down to the
level of time of day, and how these barriers function to manage
pollinators, or select for a particular pollinator guild, are as much, if
not more, a barrier than simple unrelatedness. If ever an example was needed
of the role of pollinator guild niche selection, the orchids of the
Stanhopineae contain numerous examples.

Very best

Peter

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