BLADES DEEPLY LOBED OR DIVIDED
1a. Blades deeply pinnnately lobed,
the divisions extending at least 1/4 of the way to the midrib.
2a. Leaves bipinnately
lobed, the blade divisions near the middle of the blade pinnately lobed,
3a. Blades lobed only 4/10 to
7/10 the way to the midrib; Mexico (Sinaloa & Nayarit); 0-370 m.
dressleri G.S. Bunting
3b. Blade deeply lobed, the
divisions extending more than 7/10 of the way to the midrib.
4a. Blades deciduous during
the dry season; leaf segments thin, drying papyraceous, pale yellow-green
to yellow-brown, with the minor veins distinctly visible, darker than surface;
lateral segements with usually 3 or more lobes per side, extending to below
the middle of the pinnae; Mexico (Jalisco & Chiapas) to El Salvador,
Guatemala, Honduras and Nicaragua, 300-1,500 m.
warszewiczii K. Koch & Bouché
4b. Blades never deciduous
during the dry season; leaf blade segment subcoriaceous, drying dark brown
or sometimes reddish brown, the minor veins not distinctly visible, on
drying, not markedly darker than surface; lateral segments with usually
1-2 lobes per side, usually restricted to near the apex of the pinnae;
Mexico to Colombia but in Mexico on the Caribbean slope, 0-500, rarely
to 1,860 m.
2b. Leaves merely pinnately
lobed, the blade divisions near the middle of the blade, not bipinnately
lobed, divided less than 1/2 the way to the midrib.
radiatum Schott var. radiatum
radiatum Schott, var. pseudoradiatum Croat
5a. Blade with the anterior lobe
less than 1.7 times longer than the lateral lobes; lateral lobes directed more
or less toward the apex, roughly paralleling the median lobe.
6a. Lateral lobes of the blade
broadly confluent with the medial lobe by at least 2 cm from the base of the
blade (as measured from the depths of the incised area and the apex of the
petiole); inflorescences usually 3-9 per axil.
7a. Medial lobe of the blade
with 5-12 pairs of primary lateral veins; Honduras to Panama, usually below
than 1,000 m.
1b. Blades deeply 3-lobed or trisect.
rothschuhianum (Engl.) Croat & Grayum
7b. Medial lobe of the blade
with 18-19 pairs of primary lateral veins; Costa Rica (Puntarenas, San
José), 1,000-1,250 m.
cotobrusense Croat & Grayum
6b. Lateral lobes of the blade
not confluent with or only weakly confluent with the medial lobe to no more
than 1 cm from the base of the blade; inflorescences usually one, rarely
to 3 per axil.
8a. Median lobe of blades
with more than 18 pairs of primary lateral veins, elliptic; central
Panama, 350-450 m.
8b. Median lobe of blade
with fewer than 12 pairs of primary lateral veins, usually oblanceolate
to oblong-lanceolate; widespread species, Mexico to Ecuador.
9a. Medial lobes mostly
3-3.5 times longer than broad (rarely to 1.7 times longer than
broad); primary lateral veins of medial lobe 4-12 per side, prominently
sunken; lateral lobes directed more or less toward the apex; Mexico
to Ecuador, 0-1,300 m.
9b. Medial lobes
mostly 1.5-2 times longer than broad; primary lateral veins
mostly 2-4 rarely 5 per side, weakly sunken; lateral lobes directed
more or less outward; Mexico to Costa Rica, 250-1,800 m.
5b. Blades with the anterior lobe
more than 1.7 times longer than the lateral lobes; lateral lobes directed laterally
(spreading) or even toward the base of the petiole (reflexed).
10a. Blades sagittate, the lateral
lobes directed somewhat toward the base; Mexico to Panama and South America,
0-1,900 m P.
10b. Blades hastate, the lateral
lobes directed outward at ca. 90E angle.
11a. Lateral lobes confluent
to the medial lobes usually more than 4 cm (rarely to 3.3 cm); basal veins
8-10 per side; Honduras to Panama, 0-680 m. P.
angustilobum Croat & Grayum
11b. Lateral lobes confluent
to the medial lobe usually less than 3 cm (rarely to 8 cm); basal veins
4-7 per side; Mexico to Costa Rica, 250-1,800 m.